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Under heat stress, sHSPs (such as HSPB1/HSP27) can be phosphorylated and activated via the p38/MAPK-MK2 signaling pathway (Gallagher et al., 2024), thereby initiating autophagy to clear damaged organelles via the lysosomal pathway (Cui et al., 2025)

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We searched PubMed (MEDLINE), EMBASE, the Cochrane Central Register of Controlled Trials (CENTRAL) and Web of Science from database inception to 31 October 2021 to identify eligible randomized controlled trials (RCTs)