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Interestingly, DA may also directly inhibit mitochondrial respiration [103] and DA oxidation generates toxic quinones and ROS leading to mitochondrial dysfunction [104]

Niu's MMP-2-cleavable CPP exposes cell-penetrating peptide only after matrix degradation, enabling cascade penetration absent in diffusion-only systems, though requiring hyperthermia (40 C) adds clinical complexity [182]

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