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TEER measures the net flux of all ions (cations and anions) across the epithelium and reflects the contribution of the paracellular resistance (R para ) that reflects the resistance of the TJs, the transcellular resistance (R trans ) that reflects the resistance to ions of the apical (R api ) and basolateral (R bas ) membranes and finally the sub-epithelial resistance (R sub )

Nature (2012) 483(7390):4748